Logically, they look the same. They are functionally equivalent. (?!^)and they \Gcheck two different conditions in the same place on the line, so there is no logical difference in the order in which the conditions are checked.
Conditions:
(?!^)= "we are not at the beginning of the input line\G= "we are in the place where the previous match ended
, , ( ), . , , , \G, , "" .